![]() Reports |
|
THE NORTHERN LAKE STATES
LAKE STATES NATIONAL FORESTS
Submitted By:
George E. Host and Philip L. Polzer
Natural Resources Research Institute
University of Minnesota, Duluth
5013 Miller Trunk Highway
Duluth, MN 55811
and
David T. Cleland
North Central Experiment Station
5985 Highway K
Rhinelander, WI 54501
NRRI Technical Report NRRI/TR-95/32
Center for Water and the Environment Contribution Number 162
NRRI Geographic Information Systems Number 34
The University of Minnesota is an equal opportunity educator and employer.
Introduction
Introduction
Climate and regional physiography form the highest levels of multifactor ecological
classification systems used within the Lake States region (Bailey 1985, Barnes 1982).
While the glacial geology of the Lake States has been fairly well documented, climatic
patterns have been assessed only at relatively coarse spatial scales. Rauscher (1984) used
principal component analyses of weather station data to identify 20 macroclimatic regions
within the Lake States. While the classification gives a good picture of regional climatic
patterns, these classification units are generally too broad to be used for ecological
classification purposes; often only one or two units will occur within a National Forest.
Denton and Barnes (1987, 1988) conducted a climatic classification for the state of
Michigan, which was subsequently used for identifying landscape ecosystems within the
state (Albert et al. 1986). Even this finer-resolution level classification, however, had
problems in areas where climatic changes were relatively continuous, as in the western
Manistee National Forest (Albert et al. 1986). Little regional-scale climatic work has
been conducted in Wisconsin and Minnesota.
In conducting Ecological Classification and Inventory (EC & I) on National Forests,
as well as for regional-scale interagency planning efforts, there is a clear need for
climatic maps of finer spatial resolution, on the order of 1:250,000 to 1:500,000. These
maps should be able to distinguish variations in climate attributable to lake effects,
cold air drainage, and physiographic variables. Recently, digital data on monthly
precipitation and minimum and maximum temperatures on a 1 km2 basis have become
available. These commercial products use interpolation algorithms based on 30 year weather
data to generate gridded data files (ZedX, Inc.). They represent a formidable amount of
computer processing of information from the National Climatic Data Center, and have
undergone extensive validation testing. Data are georeferenced to a grid of 30 second
longitude x 30 second latitude, which is amenable for processing with raster-based
geographic information systems.
The objective of this study was to use this high resolution climatic data to conduct a
climatic analysis and classification of Michigan, Wisconsin, and Minnesota. Remote-sensing
based analyses were used to identify fine-scale units; these were then aggregated into
more homogeneous units based on similarity. The output products are maps and GIS coverages
of the final classification in conjunction with associate data tables; special emphasis is
placed on National Forest lands.
Methods
We identified climatic regions based on a high resolution climatic database consisting
of temperature and precipitation values interpolated over a 1 km2 grid across
the study area (Russo et al. 1993). Source data were monthly averages of minimum and
maximum temperatures and precipitation derived from 30-year (1961-1990) climatological
station summaries published by the National Climatic Data Center. Multiple regressions
were used to model climatic variables as a function of latitude, longitude and elevation;
the model was then used to predict values over the 1 km2 grid (Williams and
Liebhold 1995). The accuracy of the interpolations was evaluated by comparing model
predictions with actual station values. The average absolute differences between predicted
and observed minimum and maximum temperatures were 0.56 and 0.72 C, respectively; the
average absolute difference for precipitation was 0.70 cm (ZedX, Inc., 1995). The 36
climatic data layers (12 months x 3 data types) were incorporated into ARC/INFO, a
polygon-based GIS (ESRI 1992), as point coverages, and subsequently converted into raster
format using ERDAS image processing software (ERDAS 1991).
The 36 data layers were subjected to univariate and multivariate analysis to identify
climatically homogeneous regions based on similarities in seasonal temperature and
precipitation profiles. Principal component analyses (PCA) were conducted on each of the
three data sets, and eigenvalues analyzed to identify those months that contributed most
to the overall variance of the data set. These variables were carried on to further
analyses. We also conducted PCA on a 15 layer data set using representative months
stratified across the growing season: January, March, June, August, and October. We used
five rather than twelve months to reduce the strong temporal autocorrelations between
months (Briggs and Lemin 1992). These ordination procedures allowed us to identify
variables important across the entire data set, and within each individual data set.
Variables identified in the PCA analysis were the inputs to a 15-band ERDAS iterative
clustering algorithm. This analysis assigns a cluster for each pixel based on minimum
spectral distance and then iteratively recalculates the means of each cluster until the
means no longer shift (Tou and Gonzalez 1974, ERDAS 1992). The number of classes
identified in this cluster analysis was user defined; we set an initial cluster size of
100, resulting in roughly an order of magnitude greater intensity than previous
classifications. The output classes were then aggregated into larger, more homogeneous
classes based on similarities within the dendrogram; 23 larger units were identified. Mean
values of each class for each data set used in the final analysis were calculated and
reported in tabular and map format.
Principal Component Analysis
In the principal component analysis of 12 months of 30 year average minimum
temperatures, 89% of the overall variance was loaded onto the first axis. As might be
expected, winter temperatures showed the greatest degree of variation across the Lake
States, with December, January, and February receiving loadings of 0.45, 0.45 and 0.39,
respectively. On the second PCA axis, which accounted for 10% of the overall variance, May
through July minima received high positive loadings. The distribution of February minimum
temperatures across the Lake States is shown in Figure 1. The coldest temperatures are
distributed along the northern border of Minnesota; the Superior National Forest
encompasses much of this cold region. In northern Wisconsin and Michigan, Lake Superior
moderates these colder temperatures. Only a small portion of the Ottawa NF receives this
moderating effect, however, much of the forest is in the colder interior of the Upper
Peninsula. The northern districts of the Chequamegon experience similar temperature
effects, but the southern unit and much of the Nicolet receive some moderating effects due
to latitude. Similarly, the Hiawatha has relatively warm winter temperatures due to the
moderating effects of two lakes. The strong moderating effect of Lake Michigan is evident
on the Manistee National Forest, where a gradual climatic gradient extends from the Lake
to the interior highlands. The prevailing wind patterns, coupled with the large outwash
surfaces in western Michigan result in a less steep climatic gradient compared with Lake
Michigan's effect in eastern Wisconsin. The distribution of minimum temperatures is
clearly one of the most important factors controlling the distribution of species in the
Lake States.

Figure 1. Minimum February temperatures interpolated across Michigan, Wisconsin, and
Minnesota.
A similar pattern was found with maximum temperatures. Variation in maximum
temperatures is greatest during the winter months; December and January were heavily
weighted on the first axis, which accounted for 85% of the overall variance. The second
axis, accounting for 13% of total variance, was driven by May through August temperature
maxima, with August receiving the highest eigenvalue. In contrast to minimum temperatures,
the gradient of maximum temperatures has a strong SW-NE component, with hottest August
temperatures in the continental interior (Figure 2). The prairie-forest transition of
western Minnesota is well-defined by maximum temperatures, as is the 'tension zone'
through the three states.

Figure 2. Maximum August temperatures interpolated across Michigan, Wisconsin, and Minnesota.
Seasonal variation in 30 year average precipitation was driven by the amounts of
precipitation in November, December and March. The first axis accounted for 77% of the
overall variation. November precipitation showed a strong gradation from NW to SE, ranging
from 0.5 inches in northwestern Minnesota to 2.8 inches in SE Michigan (Figure 3). The
strong elevational controls found in the temperature data were not evident in the patterns
of precipitation. Precipitation in May, June and July were the most heavily weighted
variables on the second PCA axis. This axis accounted for 13% of the overall variation,
but is likely more biologically significant than the first axis.

Figure 3. November precipitation interpolated across Michigan, Wisconsin, and
Minnesota.
Cluster Analysis
The full suite of variables selected from PCA as inputs to the classification are
presented in Table 1. Similarities among pixels were calculated based on these variables,
and an agglomerative classification performed. The study region was classified into 95
units as a function of this similarity matrix (5 of the 100 classification units specified
in the original analysis had no pixels assigned to them and were dropped).
Table 1. Variables receiving extreme eigenvalues in each data set.
Maximum Temperature |
Minimum Temperature |
Precipitation |
January |
January |
March |
February |
February |
May |
May |
May |
July |
August |
June |
November |
December |
December |
December |
The high resolution classification integrates the numerous gradients defined above.
This categorical representation of continuous data delineates the latitudinal gradient of
minimum temperatures, the SW-NE gradient of maximum temperatures, and the SE-NW gradient
of precipitation (Figure 4). Evident in this classification are the prairie-forest
transition, the tension zone, and to some degree the southern boreal forest transition
zone. Units range from relatively large, homogeneous units in the peatlands of northern
Minnesota to smaller units in the highly dissected terrain of Wisconsin's driftless area.
Since climatic models were developed independently for different states, there is some
degree of discontinuity at state boundaries. These discontinuities, however, are within
error bounds calculated for the data (J. Russo, personal communication).

Figure 4. High resolution classification of the Lake States based on PCA derived input
variables within an isodata clustering routine.
For comparison with other classification systems, the high resolution classification
was aggregated based on similarity among units. An agglomerative clustering routine was
used to combine similar units. Logical cut levels were selected based on the aggregation
patterns shown in the clustering dendrogram. Twenty-three aggregated climatic regions were
identified (Figure 5).

Figure 5. Aggregation of high resolution classification into mesoclimatic regions for
Michigan, Wisconsin, and Minnesota.
Summary
Climate is one of the main factors controlling both forest productivity and regional
species range distributions. It is also a major hierarchical level in all EC & I
systems in Region 9 (Cleland et al. 1992), as well as in analogous systems being developed
at the state level. Yet, because of difficulties in synthesizing and analyzing
information, it is one of the most poorly quantified of ECS data layers. A high-resolution
climatic classification for the northern Lake States represents a valuable asset for
specialists working in the development and interpretation of ecological units. In a
related study, for example, we used this climatic database to develop a detailed climatic
classification of the area in and around the Chequamegon National Forest (Host et al. 1996).
Other studies have used this information to predict susceptibility of sites to insect
defoliation (Williams and Liebhold 1995).
The ordinations and classifications described here are based on monthly thirty-year averages. As a result, there are several aspects of climate important to plant growth, such as growing season, that were not represented in the input data. This current classification thus represents climatic regions based on thirty years of record, and should be considered an initial high resolution stratification of the Lake States. More detailed analyses, incorporating AVHRR data, greenness indices, and other components of the weather record can be superimposed on this framework to conduct more specific, localized analyses.
Ecological classifications are the upper levels of the hierarchy are important for
addressing research questions which extend beyond National Forest boundaries. Included in
these are the role of the Forests in multi-agency regional landscape management, the
assessment of biodiversity and ecological risk at regional scales, and the identification
of ecologically-sensitive areas in the face of changing climatic conditions. Higher order
classifications provide a context within which local variation in ecosystem properties may
be assessed and quantified. This particular work supports the Lake States initiatives of
subsectional classifications, off-Forest Landtype Association development, and the
development of a framework for assessing the ecological health of Lake States forests.
Albert, D. A., S. A. Denton, and B. V. Barnes. 1986. Regional landscape ecosystems of
Michigan. School of Natural Resources, University of Michigan, Ann Arbor. 32 p.
Bailey, R. G. 1985. The factor of scale in ecosystem mapping. Envir. Manag. 9:271276.
Barnes, B. V., K. S. Pregitzer, T. A. Spies, and V. H. Spooner. 1982. Ecological forest
site classification. J. For. 80:493498.
Briggs, R. D. and R. C. Lemin, Jr. 1992. Delineation of climatic regions in Maine.
Canadian Journal of Forest Research 22:801811.
Cleland, D. T., T. R. Crow, P. E. Avers, and J. R. Probst. 1992. Principles of land
stratification for delineating ecosystems. Taking an ecological approach to management.
USDA Forest Service Watershed and Air Management Publication WO-WSA-3:40-50.
Denton, S. R. and B. V. Barnes. 1987. Spatial distribution of ecologically applicable
climatic statistics in Michigan. Can. J. of For. Res. 17:598612.
Denton, S.R. and B.V. Barnes. 1988. An ecological climate classification of Michigan: a
quantitative approach. For. Sci. 34:119-138.
ERDAS. 1992. ERDAS field guide. Version 7.5. ERDAS, Atlanta, Georgia, USA.
ESRI. 1992. Arc/Info User's Manual. Environmental Systems Research Institute, Redlands,
California, USA.
Host, G. E., P. Polzer, D. J. Mladenoff, M. A. White, and T. R. Crow. 1996. A
quantitative approach to developing regional ecosystem classifications. Ecological
Applications 6:608-618.
Rauscher, H. M. 1984. Homogeneous macroclimatic zones of the Lake States. USDA Forest
Service Research Paper NC240 39 p.
Tou, J. T., and R. C. Gonzalez. 1974. Pattern recognition principles. AddisonWesley,
Reading, Massachusetts, USA.
Williams, D. W., and A. M. Leibhold. 1995. Forest defoliators and climate change:
potential changes in spatial distribution of outbreaks of western spruce budworm
(Lepidoptera: Tortricidae) and gypsy moth (Lepidoptera: Lymantriidae). Environmental
Entomology 24:19.
ZedX. 1995. Database description: Minnesota, Michigan, and Wisconsin. HiRez Data Climatological Series, ZedX, Boalsburg, Pennsylvania, USA.